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+---------SUBJ:V-N--------+ +----COMP:N-N(of)---+ +-----COMP:V-N(in)----+ | +MOD_AT+ | +MOD_ATT:N-N+ | | | | | | Expression of __NODE__ mRNA occurs in artery from __SP__ tail . COMP:N-N(of) (expression,mRNA) MOD_ATT:N-ADJ (mRNA,__NODE__) SUBJ:V-N (occur,expression) COMP:V-N(in) (occur,__SP__) MOD_ATT:N-N (__SP__,artery)