+----------------SUBJ:V-N----------------+                              
      +-----------COMP:N-N(of)-----------+     |                              
      |           +-----MOD_ATT:N-ADJ----+     +-------COMP:V-N(in)------+    
      |           |       +--MOD_ATT:N-N-+     |         +-MOD_ATT:N-ADJ-+    
      |           |       |      +MOD_ATT+     |         |       +MOD_ATT+    
      |           |       |      |       |     |         |       |       |    
 Expression of __SP__ Igf 1 [__NODE__] mRNA occurs in __SP__ skeletal muscle .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-N (mRNA,1)
MOD_ATT:N-N (mRNA,[__NODE__])
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,muscle)
MOD_ATT:N-ADJ (muscle,__SP__)
MOD_ATT:N-ADJ (muscle,skeletal)