+--------------SUBJ:V-N-------------+                    
      +---------COMP:N-N(of)--------+     |                    
      |           +--MOD_ATT:N-ADJ--+     |                    
      |           |       +MOD_ATT:N+     +--COMP:V-N(in)-+    
      |           |       |     +MOD+     |         +MOD_A+    
      |           |       |     |   |     |         |     |    
 Expression of __SP__ __NODE__ E1 mRNA occurs in __SP__ liver .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
MOD_ATT:N-N (mRNA,E1)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,liver)
MOD_ATT:N-ADJ (liver,__SP__)