+------------SUBJ:V-N------------+                       
      +-------COMP:N-N(of)-------+     +---COMP:V-N(in)---+    
      |           +-MOD_ATT:N-ADJ+     |        +MOD_ATT:N+    
      |           |       +MOD_AT+     |        |    +MOD_+    
      |           |       |      |     |        |    |    |    
 Expression of __SP__ __NODE__ mRNA occurs in alpha TSH cells .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,cell)
MOD_ATT:N-N (cell,alpha)
MOD_ATT:N-N (cell,TSH)