+------------SUBJ:V-N------------+----------COMP:V-N(in)----------+    
      +-------COMP:N-N(of)-------+     |        +------MOD_ATT:N-N------+    
      |           +-MOD_ATT:N-ADJ+     |        |          +MOD_ATT:N-AD+    
      |           |       +MOD_AT+     |        |          |      +MOD_A+    
      |           |       |      |     |        |          |      |     |    
 Expression of __SP__ __NODE__ mRNA occurs in bone from mature __SP__ tibia .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,tibia)
MOD_ATT:N-N (tibia,bone)
MOD_ATT:N-ADJ (tibia,mature)
MOD_ATT:N-ADJ (tibia,__SP__)