+--------------COMP:V-N(in)-------------+    
      +------------SUBJ:V-N------------+        +----------MOD_ATT:N-N---------+    
      +-------COMP:N-N(of)-------+     |        |           +----MOD_ATT:N-N---+    
      |           +-MOD_ATT:N-ADJ+     |        |           |        +MOD_ATT:N+    
      |           |       +MOD_AT+     |        |           |        |   +MOD_A+    
      |           |       |      |     |        |           |        |   |     |    
 Expression of __SP__ __NODE__ mRNA occurs in colon from female 73 year old __SP__ .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,__SP__)
MOD_ATT:N-N (__SP__,colon)
MOD_ATT:N-N (__SP__,female)
MOD_ATT:N-N (__SP__,year)
MOD_ATT:N-ADJ (__SP__,old)