+------------SUBJ:V-N------------+-----------COMP:V-N(in)----------+                   
      +-------COMP:N-N(of)-------+     |         +------MOD_ATT:N-N------+                   
      |           +-MOD_ATT:N-ADJ+     |         |      +---MOD_ATT:N-N--+                   
      |           |       +MOD_AT+     |         |      |    +MOD_ATT:N-N+                   
      |           |       |      |     |         |      |    |           |                   
 Expression of __SP__ __NODE__ mRNA occurs in granule cell layer from __SP__ olfactory bulb .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,__SP__)
MOD_ATT:N-N (__SP__,granule)
MOD_ATT:N-N (__SP__,cell)
MOD_ATT:N-N (__SP__,layer)