+------------SUBJ:V-N------------+                                    
      +-------COMP:N-N(of)-------+     +---------COMP:V-N(in)---------+     
      |           +-MOD_ATT:N-ADJ+     |         +----MOD_ATT:N-ADJ---+     
      |           |       +MOD_AT+     |         |         +MOD_ATT:N-+     
      |           |       |      |     |         |         |          |     
 Expression of __SP__ __NODE__ mRNA occurs in __SP__ subcutaneous __NODE__ .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,__NODE__)
MOD_ATT:N-ADJ (__NODE__,__SP__)
MOD_ATT:N-ADJ (__NODE__,subcutaneous)