+--------------SUBJ:V-N-------------+                   
      +--------COMP:N-N(of)-------+       |                   
      |           +-MOD_ATT:N-ADJ-+       +-COMP:V-N(in)-+    
      |           |       +MOD_ATT+       |        +MOD_A+    
      |           |       |       |       |        |     |    
 Expression of __SP__ __NODE__ mRNA(s) occurs in SHEP2 cells .
COMP:N-N(of) (expression,mRNA(s))
MOD_ATT:N-ADJ (mRNA(s),__SP__)
MOD_ATT:N-ADJ (mRNA(s),__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,cell)
MOD_ATT:N-N (cell,SHEP2)