+----------COMP:V-N(in)----------+    
      +----------SUBJ:V-N---------+          +----MOD_ATT:N-ADJ----+    
      +----COMP:N-N(of)---+       |          |       +MOD_ATT:N-ADJ+    
      |           +MOD_ATT+       |          |       |      +MOD_AT+    
      |           |       |       |          |       |      |      |    
 Expression of __SP__ __NODE__ occurs in cultured __SP__ cumulus cells .
COMP:N-N(of) (expression,__NODE__)
MOD_ATT:N-ADJ (__NODE__,__SP__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,cell)
MOD_ATT:N-ADJ (cell,cultured)
MOD_ATT:N-ADJ (cell,__SP__)
MOD_ATT:N-N (cell,cumulus)