+------------COMP:V-N(in)------------+    
      +----------SUBJ:V-N---------+         +-------MOD_ATT:N-ADJ------+    
      +----COMP:N-N(of)---+       |         |       +---MOD_ATT:N-ADJ--+    
      |           +MOD_ATT+       |         |       |         +MOD_ATT:+    
      |           |       |       |         |       |         |        |    
 Expression of __SP__ __NODE__ occurs in __SP__ proximal intestinal mucosa .
COMP:N-N(of) (expression,__NODE__)
MOD_ATT:N-ADJ (__NODE__,__SP__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,mucosa)
MOD_ATT:N-ADJ (mucosa,__SP__)
MOD_ATT:N-ADJ (mucosa,proximate)
MOD_ATT:N-ADJ (mucosa,intestinal)