+--------------SUBJ:V-N-------------+                 
      +-------COMP:N-N(of)-------+        |                 
      |           +-MOD_ATT:N-ADJ+        |                 
      |           |       +MOD_AT+        |                 
      |           |       |      |        |                 
 Expression of __SP__ Rld 1 [__NODE__] occurs in intestine .
COMP:N-N(of) (expression,[__NODE__])
MOD_ATT:N-ADJ ([__NODE__],__SP__)
MOD_ATT:N-N ([__NODE__],1)
SUBJ:V-N (occur,expression)