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+-------------COMP:N-N(of)------------+ +-----------COMP:V-N(In)----------+ | +------MOD_ATT:N-ADJ------+ | +-MOD_ATT:N-ADJ-+ | | | +--MOD_ATT:N-ADJ--+ | | +MOD_ATT+SUBJ:V-N+--OBJ:V-N-+ | | +MOD_ATT+ | | | | | | | | | | In muscle , __SP__ __NODE__ protein increases expression of __SP__ __NODE__ [__NODE__] mRNA . MOD_ATT:N-ADJ (protein,__SP__) MOD_ATT:N-ADJ (protein,__NODE__) COMP:V-N(In) (increase,muscle) SUBJ:V-N (increase,protein) OBJ:V-N (increase,expression) COMP:N-N(of) (expression,mRNA) MOD_ATT:N-ADJ (mRNA,__SP__) MOD_ATT:N-ADJ (mRNA,__NODE__) MOD_ATT:N-N (mRNA,[__NODE__])