+-------COMP:N-N(of)-------+   
       +-MOD_ATT:N-ADJ-+--COMP:V_PASS-N(In)--+            |           +-MOD_ATT:N-ADJ+   
       |       +MOD_ATT+         +SUBJ:V_PASS+COMP:V_PASS-+           |       +MOD_AT+   
       |       |       |         |           |            |           |       |      |   
 In __SP__ granulosa cells , __NODE__ is involved in expression of __SP__ __NODE__ mRNA .
MOD_ATT:N-ADJ (cell,__SP__)
MOD_ATT:N-N (cell,granulosa)
COMP:V_PASS-N(In) (involve,cell)
SUBJ:V_PASS-N (involve,__NODE__)
COMP:V_PASS-N(in) (involve,expression)
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)