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+--------COMP:N-N(of)-------+ +---COMP:V-N(In)---+ | +-MOD_ATT:N-ADJ-+ +MOD_A+ +SUBJ:V-N+--OBJ:V-N-+ | +MOD_ATT+ | | | | | | | | In __SP__ liver , __NODE__ increases expression of __SP__ __NODE__ mRNA(s) . MOD_ATT:N-ADJ (liver,__SP__) COMP:V-N(In) (increase,liver) SUBJ:V-N (increase,__NODE__) OBJ:V-N (increase,expression) COMP:N-N(of) (expression,mRNA(s)) MOD_ATT:N-ADJ (mRNA(s),__SP__) MOD_ATT:N-ADJ (mRNA(s),__NODE__)