+-SUBJ:V-N+--OBJ:V-N-+COMP:N-N(of)+     
     |         |          |            |     
 Leupeptin increases expression of __NODE__ .
SUBJ:V-N (increase,Leupeptin)
OBJ:V-N (increase,expression)
COMP:N-N(of) (expression,__NODE__)