+-----------COMP:N-N(of)-----------+   
                               |           +-----MOD_ATT:N-ADJ----+   
                               |           |       +--MOD_ATT:N-N-+   
    +MOD_ATT+SUBJ:V-N+-OBJ:V-N-+           |       |      +MOD_ATT+   
    |       |        |         |           |       |      |       |   
 __SP__ __NODE__ mediates expression of __SP__ Igf 1 [__NODE__] mRNA .
MOD_ATT:N-ADJ (__NODE__,__SP__)
SUBJ:V-N (mediate,__NODE__)
OBJ:V-N (mediate,expression)
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-N (mRNA,1)
MOD_ATT:N-N (mRNA,[__NODE__])