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+---------------SUBJ:V-N---------------+ +----------COMP:N-N(in)---------+ | +COMP:N-N(of)+ +MOD_ATT:+ | | | | | | Expression of IGF 1 in mononuclear cells occurs . COMP:N-N(of) (expression,1) COMP:N-N(in) (expression,cell) MOD_ATT:N-ADJ (cell,mononuclear) SUBJ:V-N (occur,expression)