+-----------------SUBJ:V-N----------------+    
      +-------COMP:N-N(in)-------+              |    
      +COMP:N-N(of)+      +MOD_AT+              |    
      |            |      |      |              |    
 Expression of IGF 1 in organ culture system occurs .
COMP:N-N(of) (expression,1)
COMP:N-N(in) (expression,culture)
MOD_ATT:N-N (culture,organ)
SUBJ:V-N (occur,expression)