+-------------SUBJ:V-N-------------+    
      +----COMP:N-N(in)----+             |    
      +COMP:N-N(of)+       |             |    
      |            |       |             |    
 Expression of IGF 1 in __SP__ uterus occurs .
COMP:N-N(of) (expression,1)
COMP:N-N(in) (expression,__SP__)
SUBJ:V-N (occur,expression)