+---------------SUBJ:V-N---------------+                    
      +----------COMP:N-N(of)----------+     |                    
      |          +----MOD_ATT:N-ADJ----+     |                    
      |          |      +-MOD_ATT:N-ADJ+     +--COMP:V-N(in)-+    
      |          |      |       +MOD_AT+     |         +MOD_A+    
      |          |      |       |      |     |         |     |    
 Expression of large __SP__ __NODE__ mRNA occurs in __SP__ liver .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,large)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,liver)
MOD_ATT:N-ADJ (liver,__SP__)