+--------------COMP:V-N(in)-------------+    
      +------------SUBJ:V-N------------+          +--------MOD_ATT:N-ADJ-------+    
      +-------COMP:N-N(of)-------+     |          |       +----MOD_ATT:N-ADJ---+    
      |           +-MOD_ATT:N-ADJ+     |          |       |       +MOD_ATT:N-AD+    
      |           |       +MOD_AT+     |          |       |       |      +MOD_A+    
      |           |       |      |     |          |       |       |      |     |    
 Expression of __SP__ __NODE__ mRNA occurs in cultured primary __SP__ muscle cells .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,cell)
MOD_ATT:N-ADJ (cell,cultured)
MOD_ATT:N-ADJ (cell,primary)
MOD_ATT:N-ADJ (cell,__SP__)
MOD_ATT:N-N (cell,muscle)