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+------------SUBJ:V-N------------+------------COMP:V-N(in)------------+ +-------COMP:N-N(of)-------+ | +------MOD_ATT:N-ADJ------+ | +-MOD_ATT:N-ADJ+ | | +--MOD_ATT:N-ADJ--+ | | +MOD_AT+ | | | +MOD_ATT:N+ | | | | | | | | | Expression of __SP__ __NODE__ mRNA occurs in cultured __SP__ cortical astrocytes . COMP:N-N(of) (expression,mRNA) MOD_ATT:N-ADJ (mRNA,__SP__) MOD_ATT:N-ADJ (mRNA,__NODE__) SUBJ:V-N (occur,expression) COMP:V-N(in) (occur,astrocyte) MOD_ATT:N-ADJ (astrocyte,cultured) MOD_ATT:N-ADJ (astrocyte,__SP__) MOD_ATT:N-ADJ (astrocyte,cortical)