+------------SUBJ:V-N------------+                                      
      +-------COMP:N-N(of)-------+     +-----------COMP:V-N(in)----------+    
      |           +-MOD_ATT:N-ADJ+     |              +---MOD_ATT:N-ADJ--+    
      |           |       +MOD_AT+     |              |            +MOD_A+    
      |           |       |      |     |              |            |     |    
 Expression of __SP__ __NODE__ mRNA occurs in __NODE__/__NODE__ __SP__ liver .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,liver)
MOD_ATT:N-ADJ (liver,__NODE__/__NODE__)
MOD_ATT:N-ADJ (liver,__SP__)