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+------------SUBJ:V-N------------+-----------COMP:V-N(in)----------+ +-------COMP:N-N(of)-------+ | +-----MOD_ATT:N-ADJ----+ | +-MOD_ATT:N-ADJ+ | | +-MOD_ATT:N-ADJ+ | | +MOD_AT+ | | | +MOD_ATT+ | | | | | | | | | Expression of __SP__ __NODE__ mRNA occurs in __NODE__ __SP__ adipose tissue . COMP:N-N(of) (expression,mRNA) MOD_ATT:N-ADJ (mRNA,__SP__) MOD_ATT:N-ADJ (mRNA,__NODE__) SUBJ:V-N (occur,expression) COMP:V-N(in) (occur,toilet) MOD_ATT:N-ADJ (toilet,__NODE__) MOD_ATT:N-ADJ (toilet,__SP__) MOD_ATT:N-N (toilet,adipose)