+------------SUBJ:V-N------------+----------COMP:V-N(in)---------+      
      +-------COMP:N-N(of)-------+     |         +----MOD_ATT:N-ADJ----+      
      |           +-MOD_ATT:N-ADJ+     |         |      +--MOD_ATT:N-N-+      
      |           |       +MOD_AT+     |         |      |      +MOD_ATT+      
      |           |       |      |     |         |      |      |       |      
 Expression of __SP__ __NODE__ mRNA occurs in __SP__ granule cell precursors .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,precursor)
MOD_ATT:N-ADJ (precursor,__SP__)
MOD_ATT:N-N (precursor,granule)
MOD_ATT:N-N (precursor,cell)