+------------SUBJ:V-N------------+------------COMP:V-N(in)-----------+    
      +-------COMP:N-N(of)-------+     |         +------MOD_ATT:N-ADJ------+    
      |           +-MOD_ATT:N-ADJ+     |         |        +--MOD_ATT:N-ADJ-+    
      |           |       +MOD_AT+     |         |        |         +MOD_AT+    
      |           |       |      |     |         |        |         |      |    
 Expression of __SP__ __NODE__ mRNA occurs in __SP__ hippocampal dentate gyrus .
COMP:N-N(of) (expression,mRNA)
MOD_ATT:N-ADJ (mRNA,__SP__)
MOD_ATT:N-ADJ (mRNA,__NODE__)
SUBJ:V-N (occur,expression)
COMP:V-N(in) (occur,gyrus)
MOD_ATT:N-ADJ (gyrus,__SP__)
MOD_ATT:N-ADJ (gyrus,hippocampal)
MOD_ATT:N-ADJ (gyrus,dentate)