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                    +------COMP:V-N(of)-----+                
                    |       +MOD_ATT+SUBJ:V-+COMP:V-N(i+     
                    |       |       |       |          |     
 Upregulation of __SP__ __NODE__ protein occurs in synovium .
MOD_ATT:N-ADJ (protein,__NODE__)
COMP:V-N(of) (occur,__SP__)
SUBJ:V-N (occur,protein)
COMP:V-N(in) (occur,synovium)

                            +--COMP:V-N(of)-+                
                    +MOD_ATT+       +SUBJ:V-+COMP:V-N(i+     
                    |       |       |       |          |     
 Upregulation of __SP__ __NODE__ protein occurs in synovium .
MOD_ATT:N-ADJ (__NODE__,__SP__)
COMP:V-N(of) (occur,__NODE__)
SUBJ:V-N (occur,protein)
COMP:V-N(in) (occur,synovium)

       +---------MOD_ATT:N-N--------+                        
       |            +-MOD_ATT:N-ADJ-+                        
       |            |       +MOD_ATT+SUBJ:V-+COMP:V-N(i+     
       |            |       |       |       |          |     
 Upregulation of __SP__ __NODE__ protein occurs in synovium .
MOD_ATT:N-N (protein,Upregulation)
MOD_ATT:N-ADJ (protein,__SP__)
MOD_ATT:N-ADJ (protein,__NODE__)
SUBJ:V-N (occur,protein)
COMP:V-N(in) (occur,synovium)